Given the complex and multidimensional nature of human evolution, we need to develop theoretical and methodological frameworks to account for and model the dynamic feedbacks between co-operational biological and cultural evolutionary systems to better understand the processes that produced modern human behavior. Equally important is the generation of explicit theory-baed models that can be tested agains the empirical paleoanthropological record. With the aim of advancing multi-dimensional research on human biocultural evolution, we present a case study that examines evidence for culturally-driven behavioral change among Late Pleistocene hominins that altered the social niche occupied by hominins in western Eurasia, with consequences for subsequent biological and cultural evolution. We draw on a large sample of 167 Pleistocene assemblages across western Eurasia as data for this study, and employ mathematical and computational modeling to explore the feedbacks between cultural and biological inheritance. Shifts in land-use strategies changed the opportunities for social and biological interaction among Late Pleistocene hominins in western Eurasia with a cascade of consequences for cultural and biological evolution. Among these consequences were the disappearance of Neanderthals from the fossil and archaeological records, and the acceleration of cultural evolution among ancestors of modern humans.

Archaeology has an opportunity to offer major contributions to our understanding of the long-term interactions of humans and the environment. To do so, we must elucidate dynamic socioecological processes that generally operate at regional scales. However, the archaeological record is sparse, discontinuous, and static. Recent advances in computational model- ing provide the potential for creating experimental laboratories where dynamic processes can be simulated and their results compared against the archaeological record. Coupling computational modeling with the empirical record in this way can increase the rigor of our explanations while making more transparent the concepts on which they are based. We offer an example of such an experimental laboratory to study the long-term effects of varying landuse practices by subsistence farm- ers on landscapes, and compare the results with the Levantine Neolithic archaeological record. Different combinations of intensive and shifting cultivation, ovicaprid grazing, and settlement size are modeled for the Wadi Ziqlab drainage of north- ern Jordan. The results offer insight into conditions under which previously successful (and sustainable) landuse practices can pass an imperceptible threshold and lead to undesirable landscape consequences. This may also help explain long-term social, economic, and settlement changes in the Neolithic of Southwest Asia.